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$@!!!!!!!!!!!!!!!!!!!!!!!!#R#E#S#P#O#N#S#E#S!!#T#O!!#S#T#R#E#S#S(J
$@!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!(J1999,12/27 $@%+%s%@a (JBR>
OBJECTIVES
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I. INTRODUCTION
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$@!!"*9=B$$O(JFig.2.4
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$@!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!(J2000, 1/31 $@%+%s%@a (JBR>$@-6!%(JLIGHT STRESS AND PHYTOHORMONES
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$@#A!%(JLight and Indoleacetic Acid Levels
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$@!&(JMoa$@$i!J(J1989$@!K$O!"(JColeus$@$NMU$rMQ$$$F!"@V?'8w!J#R!K$,0E9u$NBP>H6h$HHf$Y$F3H;6$7$d$9$$#I#A#A$r#3G\A}2C$5$;$?$3$H!"#F#R8w$G=hM}$5$l$?MU$G$+$i$G$5$(>/NL$N#I#A#A$,F@$i$l$?$3$H$r<($7$?!J(JTable 4.4$@!K!#(J
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$@!&(JTriiodobenzoic acid$@!J#T#I#B#A!K!!#I#A#AM"AwM^@):^!!$O!"?"J*$K#R=hM}$r$7$?$H$-$G$5$(MU$NC&Mn$,A}2C$7$?$?$a$K$_$D$+$C$?!#(J
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$@!&%H%&%b%m%3%7$NCffu<4$N@.D9$O8w$K$h$C$F1F6A$r/$K4X78$,$"$k!J(Jfig.4.3$@!K!#@.0i$N:G=i$N?jB`$N$_#I#A#A$N8:>/$K4X78$9$k!#(J
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$@#B!%(JLight and Cytokinin Levels
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$@!&(JHewett and Wareing$@!J(J1973$@!K$O@V?'8w$,%]%W%i$N$/$C$D$$$F$$$k!"$b$7$/$O@Z$jN%$5$l$F$$$kMU$N#C#KNL$,A}2C$5$;$k$3$H$r8+IU$1$?!#(J
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$@!&(JQumaruddin and Tillberg$@!J(J1989$@!K$O!"#R8w$,?;?e$5$;$?(JScot pine$@!J(JPinus sylvestris$@!K$N
$@!&(JAmbler$@$i!J(J1992$@!K$O!"O72=H?1~$,0[$J$k%=%k%,%`$N#2:OG]l$G0i$F$??"J*$K$*$$$F!"#T!_(J2817$@$NLZIt=A1U$N#C#KNL$O#T!_(J7000$@$N(J1.5$@G\$K$J$C$?!#$3$NO@J8$NB?$/$N>Z5r$,!"#C#K$,?"J*$rO72=$5$;$J$$MW0x$G$"$k$H$$$&2>@b$r;Y;}$7$F$$$k!#(J
$@!&Cx/$K1F6A$5$l$K$/$/!"8w$N8:>/!J(J50$@!s8:>/!K$G#C#KG;EY$OA}$($?!J(JTable 4.5$@!K!#(J
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$@!&#C#K$N;HMQ$O?"J*$N1v%9%H%l%9$K;w$?>I>u$r<($7$?!#(J
$@!&1v%9%H%l%9$H#C#K$N4X78$rD4$Y$k$?$a$K!"(JThomas$@$i!J(J1995$@!K$O#C#K$r2aH/8=$9$k$?$a$KJQ7A$5$;$?%H%i%s%9%8%'%K%C%/!!%?%P%3$rMQ$$$?!#(J
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2000, 2/07 $@%+%s%@a (JBR>$@#C!%(JLight and GibberellinLevels
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$@!&(JPotter and Rood(1993)$@!D(JBrassica napus$@$G8w$,(J500$@&L(Jmol m-2 s-1$@$+$i(J25$@&L(Jmol m-2 s-1$@$K2<$,$k$H#G#A(J1$@ggg(J9$@g(J0$@$,A}2C$9$k$3$H$rH/8+$7$?!#(J
$@!!#G#AG;EY$,JQ2=$9$k$HCO>eIt$N?-D9$,A}2C$7$?$N$G!"$3$N$3$H$O#G#AG;EY$HCO>eIt@.D9$ND>@\E*$J4X78$r<($7$F$$$k!#(J
$@!&(JTalon and Zeevart(1990)$@!DF|D9$,#G#AG;EY$NJQ2=$r5/$3$9$3$H$r<($7$?!#(J
$@!!(JSilene armeria$@!JD9F|?"J*!K$rC;F|$+$iD9F|>r7o$K0\$9$H!"CO>eIt$H@.=OMU$G#G#A(J53$@$,8:>/$7!"#G#A(J19$@g(J0$@g$,A}2C$7!"(J $@D9F|$+$iC;F|$K0\$9$H5U$N$3$H$,5/$3$k!#(J
$@!!$3$N#G#AG;EY$NE>49$O#G#A(J53$@%*%-%7%?!<%<$,D9F|$G3h@-2=$9$k$?$a$G$"$j!"B>$N#G#A9g@.$K$*$$$F$O8w$K@)8B$5$l$F$$$kCJ3,$H$5$l$F$$$k!#(J
$@!!CO>eIt$N#G#A(J1$@$NC_@Q$O!"D9F|$*$h$S!"$b$7$/$OD9F|$,M6F3$9$k#G#A(J1$@$KBP$9$k46eIt$N@.D9$,A}2C$9$k$?$a$G$"$k!#(J
$@!&(JTalon et al.(1991)$@!D(JSpinacia oleracea$@$G8w$K$h$C$F#G#A(J53$@g(J4$@g(J9$@g(J0$@g$rA}2C$7!"0E>r7o2<$G$O8:>/$9$k$3$H$r<($7$?!#(J
$@!!8w$K$h$k#G#A(J53$@g(J0$@%*%-%7%@!<%<$N3h@-D4@a$O!"6/8w$K$h$C$F#G#ABe
$@!&(JSponsel(1986)$@!Dbd@-!J#D!K$H>f$N9b$$!J#T!K(JPea$@$rMQ$$$F!"0E>r7o2<$G@.0i$7$??"J*$HHf$Y$F#R8w$,N>J}$N$@!!#D%?%$%W$O0E>r7o2<$G#T%?%$%W$HF1$8$/$i$$@.D9$7!"#R8w$H0E>r7o2<$G#D%?%$%W$,#G#A(J1$@$r$b$C$F$$$?$N$G!"0E>r7o$G$N#D%?%$%W$N@.D9$O#G#A(J1$@$,860x$G$O$J$$!#(J
$@!&(JRoss et al.(1992)$@!Dbd@-!J#D!K$H>f$N9b$$!J#T!K(JLathyrus odoratus$@$rMQ$$$F!"#D$G$O35$7$F#G#AG;EY$ODc$+$C$?$,!"8w$,N>r7o$O!"#G#A(J1$@$r2p:_$7$?@.D9$r9b$a$k$3$H$O$J$$$H7kO@$E$1$i$l$?!#(J
$@!&(JWeller et al.(1994)$@!D%U%#%H%/%m%`$r7gB;!"#G#A(J1$@$r2a>j@8;:$9$k(JPea$@$NJQ0[BN$rMQ$$$F!"%U%#%H%/%m%`$O#G#A(J1$@G;EY$K$h$C$FCO>eIt$N?-D9$rJQ2=$5$;$J$$$3$H!"2+2=$d%U%#%H%/%m%`$,#G#A(J1$@$KBP$9$kLn@-$@!!#G#A(J19$@G;EY$O8w$,$"$?$C$F$$$kCO>eIt$ND:E@$G8:>/!"#G#A(J20$@g(J9$@gg$OA}2C$7$?!#(J
$@!&(JBown et al.(1975)$@g/(Jeskovic and Sjaus(1974)$@!D(JPhaseolus coccineus$@$H(JPea$@$N#G#A$N$h$&$J3h@-$,!"Gr?'$*$h$S@V?'8w$G8:>/$9$k$3$H$r<($7$?!#(J
$@!&(JToyomasu et al.(1992)$@!D#4F|4V$N0E>r7o$N$"$H#1F|4V8w$r$"$F$F0i$F$?(JLettus$@$Nr7o$KCV$$$?$b$N$h$j!"#G#A(J1$@$ODcG;EY!"#G#A(J19$@g(J0$@$O9bG;EY!"4^$s$G$$$k$3$H$r<($7$?!#(J
$@!!;w$?798~$,!"fu<4$N>e$H2<$rHf$Y$?$H$-$KF@$i$l$?!#$7$+$7!"8w$G=hM}$7$?;RMU$O#G#A(J19$@g$,DcG;EY$K$J$j!"#G#A(J20$@$O9bG;EY$K$J$C$?!#$3$l$O!"(JLettus$@$G$O#G#A(J1$@$,fu<4$N?-D9$rD4@a$7!"#G#A(J1$@G;EY$O8w$K$h$C$FD4@a$5$l!"8w$O#G#A(J1$@$rD4@a$9$k$3$H$K$h$C$Ffu<4$N?-D9$rD4@a$7$F$$$k$3$H$r7kO@$E$1$?!#(J
$@!&(JTable4.6$@!D?teIt?-D9$KBP$9$k8wNL$H8w$@!!;vNc$NB?$/$O8w$,#G#AG;EY$rA}2C$9$k$3$H$r<($?!#(J
$@!!I=$K<($7$?$$$/$D$+$N#G#A$O!"$[$+$N#G#A$KBP$7$FA06nJ*49B.EY$,0[$J$k7k2L!"/$9$k$3$H$O6C$/$3$H$G$O$J$$!#(J
$@!!#G#A(J1$@$,CO>eIt?-D9$ND4@a$K=EMW$G$"$m$&$H$$$&9M$($r9MN8$9$k$H!"B?$/$N;vNc$G8w$dD9F|4D6-2<$G#G#A(J1$@$,A}2C$9$k$3$H$O6C$/$3$H$G$"$k!#(J
$@#D!%(JLight and Abscisic Acid levels
$@!&8w$N6/$5!";}B3;~4V!"
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$@!!(JMohandass and Natarajaratram(1988)$@!D%$%s%I$G8w$KBQ@-$*$h$S46$@!!B@M[8w$h$j(J50$@!s8:>/$5$;$?J|J}$N2V=x$N#A#B#AG;EY$,A}2C$7$?!J46/$b$7$/$OJQ$o$i$J$+$C$?!#2V=x$N9bG;EY$N#A#B#A$O!"0zMQ$7$?%G!<%?$K4p$E$$$FA}2C$7$?8w9g@.$NF12=$K4X78$7$F$$$k$3$H$,<($5$l$?!#(J
$@!&(JBasiony et al.(1993)$@!D9b$$EZ>m?eJ,!J(J0.07MPa$@$^$G4%Ag$7!"K0OB>uBV$^$G:F?eOB!K(J
$@!!!!!!!!!!!!!!!!!!!!!!!!Dc$$EZ>m?eJ,!J(J0.015MPa$@$^$G4%Ag$7!"K0OB>uBV$^$G:F?eOB!K(J
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$@!!!!!!!!!!!!!!!!!!!!!!!!$N?"J*$NMU$O8w6/EY$NA}2C$G#A#B#A$,8:>/$7$?!J(JFig.4.6$@!K!#(J
$@!&(JHenson(1983)$@!D4%Ag%9%H%l%9$K$"$k%H%&%8%s%R%($r(J0$@g(J00$@g(J50$@g(J00$@&L(Jmol m-2 s-1$@$N8wNL;R%U%i%C%/%9L)EY$K$K$7$?$H$-!"#A#B#A$,A}2C$7$?$3$H$r<($7$?!#0E>r7o$G$O#A#B#A$O8:>/$7$?!#(J
$@!!H`$i$O!"?e%9%H%l%9H?1~$K$*$$$F#A#B#AC_@Q$KBP$9$kMU$NMFNL$,8w$K$D$h$/1F6A$rM?$($i$l$k$N$G!"%9%H%l%9$N$J$$$H$3$m$G$b8w$,#A#B#A4^M-NL$K1F6A$rM?$($k$+$b$7$l$J$$$H$$$&$3$H$r7kO@$E$1$?!J(JTable4.7$@!K!#(J
$@!&(JPhillips et al.(1980)$@!DCO>eIt$N5YL2$H#A#B#A$NLr3d$N8&5f$G!"0E4|$ND9$5$O%7%+%b%"$NMU$d:,$NM7N%$*$h$S7k9g#A#B#AG;EY$K1F6A$7$J$$$3$H$r<($7$?!#(J
$@!!$7$+$7!"#F#R$,B?$$8w$O7k9g#A#B#A$rA}$d$7!"M7N%$*$h$S7k9g$7$?;@$rH/8=$7!"#A#B#A$r8:@.$5$;$k!#8w$O$^$?!"M7N%#A#B#A$K1F6A$7$J$$!#(J
$@!&(JSaugy et al.(1989)$@!D%H%&%b%m%3%7:,$N@.D9$KBP$9$k8w$N1F6A$N8&5f$G!":,$N@.D9$,Gr?'8w$GM^@)$5$l!"M7N%#A#B#A$,$3$NH?1~$K4X78$7$F$$$k$3$H$r8+IU$1$?!#$^$?!"(J
$@$3$N#A#B#A$NH/8=$,0l;~E*$G$"$j!"#4;~4V8e!"#A#B#AG;EY$,8:>/$7!":,$N@.D9$,A}2C$9$k$3$H$r<($7$?!#(J
$@!!:,$N@.D9$NF0BV$d#A#B#A$N@8;:$K4p$E$-!"#A#B#A$K2C$($FB>$NMW0x$,@.D9M^@)$N860x$G$"$k$3$H$r<($7$?!#(J
$@!&(JTucker and Mansfield(1972)$@g5(Jucker(1977)$@!D%H%^%H$H%*%J%b%_$ND:2jM%@*$KBP$9$k;RNL$N1F6A$r8&5f$7!"#F#R8w$,N>$@!!$3$NH?1~$O!"#F#R8w$r=hM}$7$??"J*$Nf~2j$N#A#B#AG;EY$NA}2C$H0lCW$7!"#R=hM}$7$?f~2j$O!"f~@8$NCO>eIt$,@.D9$;$:!"#F#R=hM}$7$J$$?"J*$G$O@.D9$7$?!#(J
$@!!$3$l$O!"%H%^%H$N#F#R=hM}$K4X$9$k9bG;EY$N#I#A#A$O!"9b$$#A#B#AG;EY$N0];}$K860x$,$"$k$+$b$7$l$J$$$3$H$r<($7$F$$$k!#(J
$@!&(JLoveys(1979)$@!D%H%^%H$N#A#B#AG;EY$KBP$9$k%9%Z%/%H%k$N/$5$;$?!J#R8w=hM}$O(J40$@!A(J90$@!s9b$$!K!#(J
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$@!&(JKraepiel et al(1994).$@!D%?%P%3$N%U%#%H%/%m%`7gB;(J(aba1)$@$*$h$S#A#B#A$r2a>j@8;:$9$kJQ0[BN(J(Pew1)$@$H(Jaba1$@$H(J pew1$@$+$i$D$/$i$l$?JQ0[BN$r$b$A$$$?
$@!&(JWilliams et al.$@!D%"%*%&%-%/%5$rMQ$$$F!"0E>r7o$,#A#B#AG;EY$rA}2C$5$;$k$3$H$r<($7$?!##A#B#A$O(JNPR1$@$H(JNPR2$@0dEA;R$rH/8=!"0lJ}%U%#%H%/%m%`$OM^@)$9$k!#(J
$@!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!(J2000, 2/21 $@%+%s%@a (JBR>$@#E!%(JLight and Ethylene Levels
$@!&%(%A%l%s$O!"?"J*$N@.D9H?1~M^@):^$H$7$FCN$i$l$F$$$F!"?"J*$NMM!9$JH?1~$KH<$&$3$H$,Js9p$5$l$F$$$k!#!DC&Mn!"3+2V!J%"%J%J%92J!K!"6~@-H?1~!"@.=O!"O72=!"%9%H%l%9H?1~!JC9?e!"=}32!"IB86!"J*M}E*1?F0!K(J
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$@!&#R8w$O#C!a#C!"#A#C#C$NIU2C!"#C!a#CA06nBN$rM^@)$7!"#C!a#C$r8:>/$5$;$k!JNc30$"$j!K!#!D(JTable4.9
$@!&(JTabele4.10$@$O!"%Y%4%K%"$N%j!<%U%G%#%9%/$N#C!a#CG;EY$KBP$9$k8w$N6/$5$HGHD9$N1F6A$r<($7$?!#:G$b9b$$8w$G#F#R$r=|$$$?A4GHD9$O!"#C!a#CG;EY$r8:>/$5$;$?!#8w$,#1&L(Jmol m-2 $@#s(J-1$@!!$K8:>/$9$k$K$D$l$F!"#F#R0J30$O#C!a#CG;EY$,A}2C!"#F#R$O8:>/$7$?$"$H!"A}$($k!#(J
$@!&B?$/$Nr7o$G8w9g@.$d8F5[B.EY$,JQ$o$k$?$a$K#C#O(J2$@G;EY$,JQ$o$k$+$i$G(J_ $@$k!#(J
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$@!&%*%>%s$N%9%H%l%9$K$h$k#C!a#CG;EY$O!"L@>r7o$GG]M\$5$l$?%@%$%:$d%H%^%H$h$j0E>r7o$G$=$l$>$l(J2.6$@G\!"(J1.9$@G\$K$J$C$?!#(J
$@!&#A#C#C$+$i#C!a#C$X$NE>49$OL@>r7o$GM^@)!"#A#C#C$H#C!a#C$O0E>r7o$GC_@Q$9$k$,#6;~4V8e$K#C!a#C$O8:>/$9$k!#(J
$@!&8w$O:,N3Ce@8$rM^@)$7!"#C!a#C@8;:$rA}$d$9!J(JTable4.11$@!K!#(JSilver$@!"#C!a#C$NF/$-$NM^@):^$O!"8:8w$7$?8w$K$"$F$?:,$N:,N3$rA}$d$7$?!#$3$l$i$N7k2L$O!":,N3Ce@8$r8w$,M^@)$9$k1F6A$O!"A}2C$7$?#C!a#C$r7PM3$7$F5/$3$k$3$H$r<($7$F$$$k!#(J
$@!&0E>r7o$N4%Ag%9%H%l%9$N%3%`%.MU$O!"0E>r7o$N%9%H%l%9$N$J$$$b$N$h$jB?$/$N#C!a#C$r@8;:$7!"$^$?!"L@>r7o$G0i$C$?0`$l$?$b$7$/$O!"%9%H%l%9$N$J$$$b$N$h$j$b#C!a#CG;EY$,9b$+$C$?!J(JFig.4.8$@!K!#L@>r7o$G0`$l$??"J*$O!"%9%H%l%9$N$J$$$b$N$h$j$o$:$+$K9b$$#C!a#CG;EY$r<($7$?!#(J
$@!&MUNp$O$^$?!"0`$l$?MU$N#C!a#C@8;:$KBP$9$k8w$N1F6A$K4X$9$kMW0x$G$"$k$3$H$,<($5$l$F$$$k!#!J(JFig.4.9)
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$@!&(JWien and Turner(1989)$@$O!"%3%7%g%&$N#3IJnapthaleneacetic acid$@!"9g@.%*!<%-%7%s$O!"$3$Ni2$NC&Mn$rM^$($?!#(J
$@!&#C!a#C@8;:$O!"0E>r7o$^$?$OC;F|>r7o2<$G(JLilium$@$Ni2$NC&Mn$rA}$d$7$?!#$3$NC&Mn$O!"(JSilver thiosulfate$@!J#C!a#C$NF/$-$rM^@)!K$ri2$KCmF~$7$?$j!"%P%k%V$KA0=hM}$9$k$3$H$GKI$2!"#C!a#C$NF/$-$rDL$7$F0E>r7o$,Cg2p$7$F$$$kC&Mn$N$5$i$J$k>Z5r$K$J$C$?!#(J
$@!&>e5-$N7k2L$HHf$Y$F!"(JDecoteau and Craker(1987)$@$O!"(Jwung bean$@$GMU$NC&Mn$KBP$7$F!"#F#R8w$,5/$3$9#C!a#C9g@.$K$D$J$,$k>Z5r$r$_$D$1$?!##A#V#G(J(aminoethoxyvinylglycine)$@!"#C!a#C@89g@.M^@):^$O!"MU$NC&Mn$KBP$9$k#F#R!?#R8w=hM}$N1F6A$OJQ$($:!"#C!a#C$KG=NO$,$J$$$3$H<($7$?!#(J
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